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of the real lizards, family Lacertidae
Acanthodactylus scutellatus audouini BOULENGER, 1918
Bons, J. (1972) -
Cette note est destinée à faire le point sur le peuplement herpétologique du Maroc. Elle comprend: — Une liste des formes effectivement présentes au Maroc, avec les affinités biogéographiques des espèces, et éventuellement les travaux récents où ces formes sont étudiées. — Des commentaires sur les espèces africaines douteuses en Europe et européennes douteuses en Afrique. — Une liste des espèces à rechercher au Maroc. — Une discussion sommaire sur la systématique de certaines espèces marocaines.
Boulenger, G.A. (1918) -
Großhans, R. (2017) -
Nouira, S. & Blanc, C.P. & Crochet, P.-A. & Frétey, T. & Geniez, P. & Ineich, I. & Massary, J.-C. de & Ohler, A. & Tlili, W. & Lescure, J. (2022) -
A new taxonomic checklist is established for the Amphibians and “Reptiles” of Tunisia. In addition to international zoological scientific names, a French scientific name is attributed to each taxon. The Tunisian herpetofauna presently contains seven species of Amphibians and 62 of “Reptiles”.
Pellegrin, J. (1927) -
Tamar, K. & Carranza, S. & Sindaco, R. & Moravec, J. & Trape, J.-F. & Meiri, S. (2016) -
Acanthodactylus lizards are among the most diverse and widespread diurnal reptiles in the arid regions spanning from North Africa across to western India. Acanthodactylus constitutes the most species-rich genus in the family Lacertidae, with over 40 recognized species inhabiting a wide variety of dry habitats. The genus has seldom undergone taxonomic revisions, and although there are a number of described species and species-groups, their boundaries as well as their interspecific relationships are largely unresolved. We constructed a multilocus phylogeny, combining data from two mitochondrial (12S, cytb) and three nuclear (MC1R, ACM4, c-mos) markers for 302 individuals belonging to 36 known species, providing the first large-scale time-calibrated molecular phylogeny of the genus. We evaluated phylogenetic relationships between and within species-groups, and assessed Acanthodactylus biogeography across its known range. Acanthodactylus cladogenesis is estimated to have originated in Africa due to vicariance and dispersal events from the Oligocene onwards. Radiation started with the separation into three clades: the Western and scutellatus clades largely distributed in North Africa, and the Eastern clade occurring mostly from Arabia to south-west Asia. Most Acanthodactylus species diverged during the Miocene, possibly as a result of regional geological instability and climatic changes. We support most of the current taxonomic classifications and phylogenetic relationships, and provide genetic validity for most species. We reveal a new distinct blanfordii species-group, suggest new phylogenetic positions (A. hardyi, A. masirae), and synonymize several species and subspecies (A. lineomaculatus, A. boskianus khattensis and A. b. nigeriensis) with their phylogenetically closely-related species. We recommend a thorough systematic revision of taxa exhibiting high levels of intraspecific variability as well as clear evidence of phylogenetic complexity such as A. guineensis, A. grandis, A. dumerilii, and A. senegalensis and the pardalis and erythrurus species-groups.
Trapé, J.-F. & Trapé, S. & Chirio, L. (2012) -
Vicente Liz, A. & Rödder, D. & Vasconcelos Goncalves, D. & Velo-Antón, G. & Tarroso, P. & Geniez, P. & Crochet, P.-A. & Carvalho, S.B. & Brito, J.C. (2022) -
Aim: Hyper-arid sandy and rocky fields rank among the least biologically diverse habitats of the desert biome, yet knowledge of local biodiversity patterns is also ex tremely poor. In the Sahara Desert, palaeoclimate oscillations affected the extent of hyper-arid habitats, but it is unclear how these dynamics determined the evolution and distribution of local specialists. Herein, we assessed cryptic diversity, diversification patterns and spatial connectivity within a Sahara-widespread group of dryland adapted lizards. Location: Sahara-Sahel ecoregions. Taxon: Acanthodactylus scutellatus species group. Methods: Inter- and intraspecific phylogenetic structure, divergence times, spatial genetic patterns and cryptic diversity were assessed using nuclear and mitochondrial loci. The effects of topography and land cover on phylogeographic structure and diversity were tested with generalized linear models. Interspecific hybridization was evaluated using 11 microsatellites across the group`s major sympatry zone, predicted based on ecological niche models. Results: Species of Acanthodactylus scutellatus group exhibit Late Miocene origins, followed by extensive intraspecific divergence throughout the Pliocene. The northern Sahara worked as a major diversification hotspot, harbouring a patchwork of small ranged, divergent lineages. These lineages are parapatric or sympatric and present concordant nuclear and mitochondrial differentiation, suggesting species status. Genetic connectivity increases in southern latitudes, with wide-ranging lineages spanning from the Red Sea to the Atlantic coast. Within these potential corridors, mountain outskirts and sand fields in the Sahara interior seemingly acted as origins for recent population expansions. Genetic diversity and connectivity are favoured by terrain roughness and soft-sand cover respectively. Three species inhabit the Atlantic Sahara sympatry zone without evidence of gene flow. Main conclusions: Overlooked species-level diversity within a major specialist group of Sahara drylands exposes the recurrent knowledge shortfalls present in hyper-arid desert environments. Humidity and sandy habitat shifts triggered potential successions of population isolation and re-connectivity, which favoured cladogenesis in northern desert regions and population expansions across southern east–west corridors.